加贺美（Kagami）拉着马丽娜（Marinette）前往东京，协助“恒心战队（Stellars）”对抗全新的超级反派。但马丽娜很快发现，好友此次日本之行，还藏有不为人知的另一层缘由。

By daylight, Marinette Dupain-Cheng and Adrien Agreste are ordinary teenagers. But unbeknownst to their family and friends, the two of them carry a huge secret...using the powers of magical creatures called Kwami, they transform into the superheroes Ladybug and Cat Noir! With a mission to protect the city of Paris from the evil Hawk Moth, the two of them must learn to cope with their new responsibilities, as well as ordinary troubles involving love, school, family, and friends. Most of all, they must learn to put their romantic feelings aside in order to perform their duty, unaware of each other's true identities.