2045 年，东京——年轻的科技巨头 ZIC 的技术开发工程师大空浩史，发明了一套能够扭曲物理定律的超级战衣，使他成为史上最强的超级英雄。他的英勇事迹很快为他赢得了“东京队长”的称号，他也因此成为媒体的宠儿。

与此同时，一群名为“玩偶”的致命但又魅力四射的传送型女性机器人，受一位自称“机械博士”的神秘邪恶科学家控制，开始了一系列离奇的犯罪活动。莫尔迪弗挫败了她们的首次盗窃企图，她们原本打算从 ZIC 的赛车皇后大赛中偷走一辆经典的 F1 赛车。

那场比赛的获胜者恰好是大学生、商场常客、哥哥浩史的妹妹大空未来。她决定要揭开这位神秘英雄的真实身份！而答案或许比她想象的还要出人意料……

A naïve young girl accidentally modifies her brother's secret superhero uniform and becomes the unwilling superheroine Moldiver.

The Boonboomgers' run in the Big Bang Grand Prix is violently interrupted when the universe's top-ranked monster, Universe No. 1, appears and triggers a catastrophic universe fusion, causing parallel worlds to merge into one chaotic continuum. While Taiya Hando/Bun Red is suddenly transported to Planet Shaft, he's attacked by the Bitter Demon Beast Galaxy Circuit Guruma until Hoeru Tono/Gozyu Wolf arrives to back him up. Meanwhile on Earth, the remaining Gozyugers battle Universe No. 1. Bun Blue, Bun Pink, and Bun Black return to help, but the monster overwhelms them and defeats the combined teams. Hoeru then reappears in a dramatic new evolution: Saint Gozyu Wolf, declaring his vow to "stop the battle." With a slash that cuts through space itself, he sends all eight heroes hurtling into another dimension, setting the stage for the film's multiversal conflict.

Long ago, the gate to the world of darkness was sealed. Now evil warriors, known as Remnants, are trying to reopen the gate and let demons take over the world. However, at Tobira High School, there exist guardians of the gate who must risk their lives to prevent the gate from being opened.